Arctotheca calendula (L.) Levyns(=Arctotis calendula L.; Cryptostemma calendula (L.) Druce)Family: Asteraceae, Tribe: Arctoteae |
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Common names: capeweedDisseminule: FruitDescriptionFruit an achene, obovate, flattened, with a mildly lobed collar at the apex, 2–4(5) mm long, 1.0–1.4 mm wide, 0.7–1.0 mm thick; cross-section elliptic with distinct ridges at the margins to rhombic with a total of 3–5 longitudinal ribs; surface dark brown, granular, rugose, pubescent with long, crinkly light brown hairs that envelop the fruit to create a ball 6–8 mm diameter. Scar basal, round. Persistent pappus of 6–8 lanceolate scales, ca. 0.8 mm long each, yellowish-purple with purple veins. Style base dark brown, cylindrical. Embryo spatulate; endosperm absent. Identification considerationsThe ball of woolly hairs encasing the fruit is distinctive. DistributionNative to southern Africa. Naturalized in Australia, New Zealand, Portugal, southwestern Spain, the Azores and California, United States. HabitatThe natural habitat includes sand dunes, stream banks and rocky outcrops. Capeweed prefers sunny sites and sandy, well-drained soils. It is a tough plant, withstanding drought conditions. As a weed, it invades pastures, yards and disturbed sites, outcompeting other vegetation. General informationArctotheca calendula is a rosette-forming perennial. Fruits are believed to be wind-borne and may become lodged in animal fur. Human activities that result in the movement of contaminated soils can spread both fruits and stolons. International dispersal is sometimes intentional, the plant being valued as an ornamental groundcover and for medicinal purposes. Unintentional spread may occur via contaminated fleece, grain or seed shipments from South Africa or Australia. |
Achene with hairs teased back to better reveal the fruit Achene with hairs and pappus scales removed Close-up of the ring of pappus scales Apex with scales still attached (left) and removed (right) Attachment scar of a fruit with hairs still attached (left) and where hairs have been removed Floral heads with the hairy achenes still attached A, Fruit; B, embryo in situ; C, transection of fruit drawing by Lynda E. Chandler |
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